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Chlorophyll fluorescence was measured using a fluorometer (Open FluorCam FC 800-O/2020, Photon Systems Instruments, Brno, Czech Republic) and F v /F m was analyzed based on the fluorescence image

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*Correspondence: Yongping Liu, [email protected] These authors have contributed equally to this work and share first authorship This article was submitted to Autoimmune and Autoinflammatory Disorders : Autoimmune Disorders, a section of the journal Frontiers in Immunology Disclaimer All claims expressed in this article are solely those of the authors and do not necessarily represent those of their affiliated organizations, or those of the publisher, the editors and the reviewers